These don't seem to have been products of Russia, though. The bots were linked to a social networking outfit that previously disseminated positive stories about Saudi Arabia, including Crown Prince Mohammed bin Salman. It's not certain if the accounts were connected to the Saudi government, although Salah Faya, the person who registered the domain for the website (Arabian Veritas) linked to the accounts, described himself in part as an "e-Government Expert" and worked for a "confidential" company in Saudi Arabia. Twitter has previously fought pro-Saudi bots and may have even had an employee who was enticed to spy on Saudi dissidents.
As Watts noted, the goal wasn't necessarily to sway people directly. They were too obviously artificial for that. Rather, it was to amplify the overall signal of the keywords and influence people inclined to believe the political message behind them. And when it's relatively simple and inexpensive to start a bot network, the results don't need to be dramatic to have value. Twitter may have a tough fight to prevent networks like this from surfacing in the future, even if it's more capable of dealing with them than it was a few years ago.